complex cell

Visual-cortex cells {complex cell} can receive from simple cells and ganglion cells. Complex cells have ocular dominance or orientation tuning.

functions

They can detect stereoscopic effects, such as line-segment ends, colors, motions, and line orientations. They can mark region boundaries, such as regions with same reflectance. They discriminate and aggregate. They can detect patterns, at any location.

fields

Complex cells have different receptive-field sizes and detect different spatial frequencies and so widths. Sets can detect 8 to 30 different frequency bands and spatial scales, using spatial frequency channels, from pixel, spot, region, quadrant, or whole visual field.

time

Complex-cell sets can operate at 10 to 30 different temporal scales, from milliseconds to years.

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