coupled fitness landscape

Gene fitness can depend on other-gene fitness in same or other species {coupled fitness landscape} {fitness landscape, coupled}.

If species dependency is low, same-species gene dependency is high, or species number is low, system is stable. If species dependency is high, same-species gene dependency is low, or species number is high, system is unstable.

If species number or species-dependency level can change, species dependency tends to middle, to maximize average fitness and minimize average extinction rate. Middle level is just before point where too many changes happen.

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